Keywords
hemerophytes
cultivated plants
flora
introduction
ergasiophytes
plant invasions
Kyiv
Cherkasy Oblast
climate change
ecological risks
How to Cite
Abstract
This study presents a comprehensive analysis of the taxonomic diversity, acclimatization processes, and naturalization success of ornamental herbaceous perennials and semi-woody plants in Central Ukraine. The role of these plants in regional floriculture and the ecological risks associated with their introduction are discussed. It was found that regional floriculture probably has a long history, but the first specific records on the study object appeared at the end of the 18th century. The research revealed that 794 species, subspecies, and hybrids from 301 genera and 70 families are cultivated in the regional floriculture. The largest number of species and infraspecific taxa belong to the families Asteraceae (11.6 %), Asparagaceae (6.5 %), Lamiaceae (6.5 %), Ranunculaceae (6.0 %), and Crassulaceae (5.3 %). The most represented genera are Allium (25 species), Iris (19 species and hybrids), and Primula (14 species and hybrids). It was found that 84.5 % of the studied species and infraspecific taxa are ergasiophytes, while 15.5 % are native plants, often represented by cultivars, reflecting the predominance of introduced species and cultivars in the assortment of ornamental plants. Among the plants used in floriculture in Central Ukraine, herbaceous perennials species constitute the largest group (77.5 %), while the presence of semi-woody plants (5.1 %) and annual and biennial plants (17.4 %) is significantly lower. The distribution of native species by range types covers all major elements of the natural flora, but species with European (23.6 %), Eurasian (19.5 %), and European-Mediterranean (13.9 %) distribution patterns are the most frequently cultivated. Among ergasiophytes, most species and infraspecies have Asian (28.0 %), Mediterranean (19.4 %), and American (19.1 %) origin, with a significant proportion of hybrids and cultigenous species (11.2 %). Overall, species from all geographic regions, including tropical and oceanic zones, are represented in floriculture.
An essential aspect of the study was assessing the acclimatization and naturalization degrees of ornamental alien plants. The scheme for overcoming limiting barriers by alien species was supplemented with a model describing the acclimatization of ergasiophytes and their escape beyond cultivated areas. The acclimatization of ergasiophytes in this study is considered a controlled process that is ongoing simultaneously with spontaneous naturalization. It was found that 44.9 % of ergasiophytes achieved complete acclimatization, 15.4 % penetrated beyond cultivation sites, becoming ergasiophygophytes, 2.7 % naturalized, and 1.5 % acquired invasive status. For example, invasive plants include Helianthus tuberosus, Reynoutria japonica, and Solidago canadensis. Potentially invasive species requiring monitoring and further study comprise Corydalis caucasica, Petrosedum orientale, Symphyotrichum × versicolor, Thladiantha dubia, and others.
References
Abduloyeva, O.S., & Karpenko, N.I. (2009). Occurrence of alien invasive plant species in vegetation syntaxa of Ukraine. Chornomorski Botanical Journal, 5(2), 189–198. (In Ukrainian)
Alekhin, A.A., Orlova, T.G., & Alekhina, N.N. (2011). The flower-decorative plants in the Botanical Garden of V.N. Karazin Kharkov National University. In Proceedings of the 3rd International Conference on Landscape Architecture in Botanical Gardens and Arboretums (pp. 136–140). Kyiv. (In Russian)
Alyokhin, A.A., Orlova, T.G., & Alyokhina, N.N. (2021, March 31). Changes in the composition of the collection of flower ornamental plants in connection with climate aridization. In Proceedings of the International Scientific Conference “Conservation of plants in connection with climate changes and biological invasions” (pp. 9–11). Bilotserkivdruk, Bila Tserkva. (In Ukrainian)
Andrukh, N.A. (2016). Genus Heuchera L. in the Right-Bank Forest-Steppe of Ukraine: biological features and introduction (PhD thesis. M.M. Gryshko National Botanical Garden of National Academy of Sciences of Ukraine). Kyiv. (In Ukrainian)
Andrzejowski, A. (1861). Enumeratio plantarum sponte in gubernio Podolico et locis adjacentibus crescentium. Works of the Commission, Supremely established at the Imperial University of St. Vladimir for the description of the provinces of the Kyiv educational district, 4(1), 1–51. (In Russian)
Antonyuk, N.Y., Borodina, R.M., Stopkan, V.V., & Skvortsova, L.S. (1977). Decorative plants of the natural flora of Ukraine. Naukova Dumka, Kyiv. (In Ukrainian)
Arkushyna, H.F., & Popova, O.M. (2010). Checklist of the flora of vascular plants of Kirovograd. Polimed-Servis, Kirovograd. (In Ukrainian)
Barbarich, A.I. (1938). Ornamental plants of the Right-Bank Polissia and ways to replenish their assortment. In M.G. Kholodny (Ed.), Collection of works dedicated to the memory of the academician O.V. Fomina (pp. 195–203). Publishing Academy of Sciences of the Ukrainian SSR, Kyiv. (In Ukrainian)
Barbarich, A.I. (1940). Decorating plants in the Donets Basin. Ukrainian Botanical Journal, 1(3–4), 353–359. (In Ukrainian)
Barbarich, A.I. (1945). Decorative plants of the Ukrainian S.S.R. Botanical Journal, 2(3–4), 159–166. (In Ukrainian)
Barbarich, A.I. (1972). Ornamental plants of the Ukrainian Polesye settlements in the XIX–first half of the XX century. Ukrainian Botanical Journal, 29(5), 662–665. (In Ukrainian)
Beauplan, G.L. (1660). Description d’Ukranie, qui sont plusieurs provinces du Royaume de Pologne, contenues depuis les confins de la Moscovie jusques aux limites de la Transilvanie: ensemble leurs moeurs, façons de vivres & de faire la guerre. Jacques Cailloüé, Rouen.
Berezkina, V. (2007). Assessment of the success of the introduction of Sedum L. species. Bulletin of Taras Shevchenko National University of Kyiv. Series: Introduction and Preservation of Plant Diversity, 11, 4–6. (In Ukrainian)
Berezkina, V.I., Kukovytsya, G.S., Menshova, V.O., Vasheka, O.V., Gadzhosa, A.Y., Yeresova, A.V., Holubenko, A.V., Rudik, H.O., & Syvets, G.V. (2007). Catalogue of herbaceous plants. In O.V. Fomin Botanical Garden: Catalog of plants (pp. 71–124). Phytosociocentre, Kyiv. (In Ukrainian)
Biliavskyi, S.M. (2021). Urban flora of Bila Tserkva town and its suburbs (PhD thesis. M.M. Gryshko National Botanical Garden of National Academy of Sciences of Ukraine). Kyiv. (In Ukrainian)
Bilous, V.I. (2001). Garden and park art: a brief history of development and methods of creating artistic gardens. Naukova Dumka, Kyiv. (In Ukrainian)
Boiko G.V. (2011). Genus Artemisia L. (Asteraceae Bercht. & J. Presl) in the flora of Ukraine. (PhD thesis. M.G. Kholodny Institute of Botany National Academy of Sciences of Ukraine). Kyiv. (In Ukrainian)
Boiko, I.V. (2024). Methodological recommendations for growing representatives of the genus Helleborus (Hellebore). Sochynskyi M.M., Uman. (In Ukrainian)
Bomble, F.W. (2016). Kultivierte und verwildernde Arten von Phedimus subgen. Aizoon im Aachener Raum und im Ruhrgebiet. Jahrbuch des Bochumer Botanischen Vereins, 7, 17–36.
Bonetskyi, S. (1927). Industrial and estate floriculture. Radyanskyi Selyanyn, Kharkiv. (In Ukrainian)
Bonnier, G., Douin, R., Poinston, J., & Palese, R. (1990). La grande flore en couleurs de Gaston Bonnier. Belin, Paris.
Brashear, C., & Meyer, B. (2025). Hosta Library. https://www.hostalibrary.org/
Brickell, C. (Ed.). (2011). Encyclopedia of plants & flowers. DK Publishing, New York.
Buidin, Y. (2004). Morphological and biological features of introduced varieties of Astilbe Buch.-Ham. ex D. Don in the Forest-Steppe of Ukraine (PhD thesis. M.M. Gryshko National Botanical Garden of National Academy of Sciences of Ukraine). Kyiv. (In Ukrainian)
Buidin, Y., & Skrypka, A. (2009, October 28–29). The collection fund of bearded irises at the M.M. Gryshko National Botanical Garden of Ukraine. In Proceedings of the 9th All-Ukrainian Scientific Conference “Biological studies of young scientists in Ukraine” (pp. 7–9). Kyiv. (In Ukrainian)
Burda, R.I. (2013). Plant introduction:domestication and naturalization. Industrial Botany, 13, 3–15. (In Russian)
Burda, R.I., Pashkevich, N.A., Boyko, G.V., & Fitsaylo, T.V. (2015). Alien species of protected flora of the Forest-Steppe of Ukraine. Naukova Dumka, Kyiv. (In Ukrainian)
Byalt, V.V., Firsov, G.A., Byalt, A.V., & Orlova, L.V. (2019). Overview of the cultural flora of St. Petersburg (Russia). Moscow. (In Russian)
Cherniak, V.M., Shehera, L.I., & Trofymenko, N.M. (1998, October 5–8). Ornamental plants of Western Podillia – Prospects for landscaping and improvement of settlements. In Materials of the 6th International Conference “Problems of dendrology, floriculture, and fruit growing” (pp. 94–97). Yalta. (In Ukrainian)
Chikov, I.V. (2016). Methodological recommendations for growing ornamental aquatic and coastal aquatic plants in the Right-Bank Forest-Steppe of Ukraine. Palyvoda A.V., Kyiv. (In Ukrainian)
Chopyk, V.I., Bortniak, M.M., Voytiuk, Y.O., Pohrebennyk, V.P., Kucheriava, L.F., Nechytailo, V.A., Liubchenko, V.M. & Shevchyk, V.L. (1998). Synopsis of the flora of the Middle Dnipro region: vascular plants. Phytosociocentre, Kyiv. (In Ukrainian)
Chorna, H.A. (2006a). Flora of the ponds and bogs of the Forest-Steppe zone of Ukraine: vascular plants. Phytosociocentre, Kyiv. (In Ukrainian)
Chorna, H.A. (2006b). Invasion of ornamental alien plants of the river valley of the forest-steppe zone of Ukraine. Biological Herald (Kharkiv), 10(2), 49–51. (In Ukrainian)
Chorna, H.A. (2020, September 22–24). Species of the genus Phytolacca L. in Ukraine. In Proceedings of the International Scientific Conference Dedicated to the 85th Anniversary of the M.M. Gryshko National Botanical Garden of NAS of Ukraine (pp. 297–301). Lira-K, Kyiv. (In Ukrainian)
Chorna, H.A., & Kostruba, T.M. (2019, September 26–27). Amateur floriculture and phytoinvasions. In Proceedings of the 3rd All-Ukrainian Scientific Conference “Synanthropization of the plant cover of Ukraine” (pp. 175–180). Nash Format, Kyiv. (In Ukrainian)
Chorna, H.A., Shynder, O.I., & Kostruba, T.M. (2021). Addition to the list of species of the spontaneous flora of the National Dendrological Park Sofiyivka of the National Academy of Sciences of Ukraine (Uman, Cherkasy Region). Chornomorski Botanical Journal, 17(4), 302–315. (In Ukrainian). https://doi.org/10.32999/ksu1990-553X/2021-17-4-1
Cullen, J., Knees, S.G., & Cubey, H.S. (2011). The European garden flora. Cambridge University Press.
Davydov, D. (2025). Biodiversity of the Left Bank Forest-Steppe of Ukraine. iNaturalist. https://www.inaturalist.org/projects/biodiversity-of-the-left-bank-forest-steppe-of-ukraine
Didenko, S.Y. (2000). Species of the genus Galanthus L. (Amaryllidaceae) in nature and culture in Ukraine (PhD thesis. M.M. Gryshko National Botanical Garden of National Academy of Sciences of Ukraine). Kyiv. (In Ukrainian)
Didenko, V.I., Kuzemko, A.A., & Shynder, O.I. (2024). Spontaneous and cultural flora of the National Scientific Center “P.I. Prokopovich Beekeeping Institute” territory (Kyiv). Chornomorski Botanical Journal, 20(2), 168–189. (In Ukrainian). https://doi.org/10.32999/ksu1990-553X/2024-20-2-4
Didenko, V.I., Kuzemko, A.A., Bezsmertna, O.O., Shynder, O.I., Kucher, O.O., Shevchyk, V.L., Podobaylo, A.V., & Kostikov, I.Y. (2022). Common milkweed (Asclepias syriaca L., Apocynaceae Juss.) – the highly invasive and honey bearer species of the Ukrainian flora. Beekeeping of Ukraine, 9, 27–39. (In Ukrainian). https://doi.org/10.46913/beekeepingjournal.2022.9.04
Didukh, N.Y., & Mazur, T.P. (2013). Representatives of the genus Nuphar Smith in natural conditions of Ukraine and culture: biological, economic, and morphological features. Publishing and Printing Center “Kyiv University”, Kyiv. (In Ukrainian)
Didukh, Y.P., & Chorney, I.I. (2016). Climatogenic changes of plant life of the Ukrainian Carpathians. DrukArt, Chernivtsi. (In Ukrainian)
Doiko, N.M., Kalashnikova, L.V., & Rubis, V.L. (2013). Catalogue of herbaceous plants of the State Dendrological Park “Oleksandria” of the National Academy of Sciences of Ukraine. Bila Tserkva. (In Ukrainian)
Doiko, N.M., Shynder, O.I., & Dragan, N.V. (2021). Regional features and long-term dynamics of the flora of the “Oleksandria” Dendrological Park of the NAS of Ukraine (Bila Tserkva). Ecological Sciences, 7(34), 81–90. (In Ukrainian). https://doi.org/10.32846/2306-9716/2021.eco.7-34.14
Dovzhenok, V.Y. (1961). Agriculture of Ancient Rus’ before the middle of the 13th century. Publishing House of the Academy of Sciences of the Ukrainian SSR, Kyiv. (In Ukrainian)
Dubniak, K. (1924). On the decorative flora of the Myrhorod city surroundings, Poltava Region. Ukrainian Botanical Journal, 2(1922), 52–54. (In Ukrainian)
Dudarets, V.M. (2019). Historical origins of garden art in Kyivan Rus. In M.I. Stepanenko (Ed.), Ethnodesign in the Context of Ukrainian National Revival and European Integration (pp. 232–238). PNU V.G. Korolenko, Poltava. (In Ukrainian)
Fedorchuk, M.I., Bazalii, V.V., Mrinskyi, I.M., Onyshchenko, S.O., Mazurok, I.H., & Kotovska, Y.S. (2012). Perennial ornamental plants of the dendrological park of Kherson State Agrarian University. Hrin D.S., Kherson. (In Ukrainian)
Fedorov, A.A. (Ed.). (1974–1987). Flora partis europaeae URSS. Vols. 1–6. Nauka, Leningrad. (In Russian)
Flora of North America Editorial Committee (Eds.). (1993–2023). Flora of North America North of Mexico. Vols. 1–28. Oxford University Press, New York. https://floranorthamerica.org/
Fritsch, R.M., & Abbasi, M. (2013). A taxonomic review of Allium subg. Melanocrommyum in Iran. Leibniz-Institut für Pflanzengenetik und Kulturpflanzenforschung, Gatersleben.
Galkin, S.I. (2013). Park “Oleksandria”: history and modernity. Pshonkivskyi O., Bila Tserkva. (In Ukrainian)
Gallo, L., & Zika, P.F. (2014). A taxonomic study of Sedum series Rupestria (Crassulaceae) naturalized in North America. Phytotaxa, 175(1), 19–28. https://doi.org/10.11646/phytotaxa.175.1.2
Glukhov, O., Prokhorova, S., Derevyanska, G., & Kharkhota, G. (2010). Ornamental alien plants of natural flora in the anthropogenic megalopolis Donetsk-Makeevka. Plant Introduction, 45, 3–9. (In Ukrainian). https://doi.org/10.5281/zenodo.2553584
Glukhova, S.A., Shynder, O.I., Mykhailyk, S.M., & Yemets, L.I. (2019). Cereals and grass-like herbs in the collection of the Syrets Arboretum of National Importance (Kyiv). News of the Biosphere Reserve “Askania Nova”, 21, 414–416. (In Ukrainian). https://doi.org/10.53904/1682-2374/2019-21/64
Glukhova, S.A., Shynder, O.I., Yemets, L.I., & Mykhailyk, S.M. (2016). Catalogue of herbaceous plants of Syretsky dendrological park. Poltavskyi Literator, Poltava. (In Ukrainian)
Golub, V., & Golub, N. (2002). Decorative hydrophytes of the Right-Bank Forest-Steppe of Ukraine and perspectives for their use in water reservoir decoration. Plant Introduction, 13, 129–132. (In Ukrainian). https://doi.org/10.5281/zenodo.3333817
Gorobets, V.F. (Ed.). (2009). Catalogue of ornamental plants for urban and suburban landscaping of the Forest-Steppe and Polesia of Ukraine. Landscape and Interiors (Kyiv), 7, 1–188. (In Russian)
Gorobets, V.F., Mashkovska, S.P., & Buydin, Y.V. (2008). Collection fund of flower-ornamental plants of M.M. Gryshko NAS Ukraine National Botanical Garden. Catalogue (Index Plantarum). Medobory, Ternopil. (In Ukrainian)
Govaerts, R. (2023). The world checklist of vascular plants (WCVP). Ver. 12. The Royal Botanic Gardens, Kew. https://doi.org/10.34885/jdh2-dr22
Grechyshkina, Y.V. (2010). Natural flora of vascular plants of Kyiv. (PhD thesis. M.G. Kholodny Institute of Botany of the National Academy of Sciences of Ukraine). Kyiv. (In Ukrainian)
Grodzinsky, A.M. (Ed.). (1985). Ornamental plants for open and closed grounds. Naukova Dumka, Kyiv. (In Russian)
Grossheim, A.A. (1949). Key to plants of the Caucasus. Soviet Science, Moscow. (In Russian)
Gubar, L M., & Koniakin, S.N. (2020). Invasive alien species of plants of the local landscape Feofania. Ecological Sciences, 4(31), 167–173. (In Ukrainian). https://doi.org/10.32846/2306-9716/2020.eco.4-31.26
Güldenstädt, J.A. (1791). Reisen durch Russland und im Caucasischen Gebürge. Th. 2. Russisch-Kayserlichen Akademie der Wissenschaften, Saint Petersburg.
Herasymiuk, N.V. (2012). Ornamental plants of the private sector in Odesa. Tavriiskyi Scientific Bulletin, 80(2), 67–71. (In Ukrainian)
Hoffmann, M.H. (1996). Die in Zentraleuropa verwilderten und kultivierten nordamerikanischen Astern. Feddes Repertorium, 107(3–4), 163–188. https://doi.org/10.1002/fedr.19961070305
Holub, J., & Jirásek, V. (1967). Zur Vereinheitlichung der Terminologie in der Phytogeographie. Folia Geobotanica et Phytotaxonomica, 2, 69–113. https://doi.org/10.1007/BF02851755
Horai, H.O. (2007). Biological features and prospects for the use of decorative species of the family Papaveraceae Juss., introduced in the Forest-Steppe of Ukraine. Proceedings of the National Natural History Museum of Ukraine. Botanical Series, 2, 355–364. (In Ukrainian)
Horbenko, N.Y., & Hrynyk, O.M. (2010). Use of shade-tolerant forest plants and their decorative forms in the landscaping of Lviv. Bulletin of the National University of Life and Environmental Sciences of Ukraine. Series: Forestry and Decorative Gardening, 152(1), 13–17. (In Ukrainian)
ICN. (2025). International Crassulaceae Network. https://www.crassulaceae.ch/
iNaturalist. (2025). iNaturalist. A community for naturalists. https://www.inaturalist.org/
Ishchuk, L.P. (2012, May 15–18). Flower-decorative landscaping of the village of Kovalyvka, Vasylkiv District, Kyiv Region. In Proceedings of the International Readings Dedicated to the 110th Anniversary of Professor L.I. Rubtsov (pp. 144–148). Molyar S.V., Kyiv. (In Russian)
Kalusová, V., Čeplová, N., Danihelka, J., Večeřa, M., Pyšek, P., Albert, A., Anastasiu, P., Biurrun, I., Boch, S., Cottaz, C., Essl, F., Kuzemko, A., Maslo, S., Mifsud, S., Protopopova, V., Shevera, M., Sîrbu, C., Svenning, J.-Ch., Welk, E., & Axmanova, I. (2024). Alien plants of Europe: an overview of national and regional inventories. Preslia, 96(2), 149–182. https://doi.org/10.23855/preslia.2024.149
Karmazin, R.V., & Lyskovych, Z.M. (1978). Ground cover perennial ornamental plants in the landscaping of Lviv region. In Biological features of useful plants of the natural flora in connection with their introduction in Ukraine (pp. 50–53). Naukova Dumka, Kyiv. (In Ukrainian)
Kharkevich, S.S. (1966). Naturalization of plants of the natural flora of the Caucasus in Kiev. Bulletin of the Main Botanical Garden, 61, 3–8. (In Russian)
Khassanov, F.O., & Fritsch, R.M. (1994). New taxa in Allium L. subg. Melanocrommyum (Webb & Berth.) Rouy from Central Asia. Linzer biologische Beiträge, 26(1), 965–990.
Kleopov, Y.D. (1990). Analysis of the flora of deciduous forests of the European part of the USSR. Naukova Dumka, Kyiv. (In Russian)
Klymenko, S., Ilyinska, A., Kalista, M., & Grygorieva, O. (2019). Aubrieta deltoidea (L.) DC. (Brassicaceae) – potential ergasiophyte of the flora of Ukraine and Eastern Europe. Plant Introduction, 82, 55–63. (In Ukrainian). https://doi.org/10.5281/zenodo.3241097
Kokhno, M.A. & Trofymenko, N.M. (2005). Dendroflora of Ukraine. Wild and cultivated trees and bushes. Angiosperms. Part 2. Phytosociocentre, Kyiv. (In Ukrainian)
Kokhno, N.A. (1983). On the assessment of the success of the introduction of woody plants. Introduction of woody plants and greening of cities in Ukraine: collection of scientific papers. Naukova Dumka, Kyiv. (In Russian)
Kokhno, N.A. (Ed.). (1997). Catalogue of plants of the M.M. Gryshko Central Botanical Garden. Naukova Dumka, Kyiv. (In Russian)
Kokhno, N.A., & Kurdyuk, A.M. (1994). Theoretical foundations and experience of the introduction of woody plants in Ukraine. Naukova Dumka, Kyiv. (In Russian)
Kolomiychuk, V., & Shynder, O. (2021). Addition to the spontaneous flora of O.V. Fomin Botanical Garden (Kyiv). Bulletin of Taras Shevchenko Kyiv National University. Series: Biology, 87(4), 18–26. (In Ukrainian)
Koniakin, S.M., Burda, R.I., & Budzhak, V.V. (2023). The alien flora of the Kyiv urban area, 2003–2022: Prelude notes. Chornomorski Botanical Journal, 19(2), 200–225. (In Ukrainian). https://doi.org/10.32999/ksu1990-553X/2023-19-2-4
Kornaś, J. (1978). Remarks on the analysis of a synantropic flora. Acta Botanica Slovaca A, 3, 385–394.
Kosenko, I.S. (Ed.). (2000). Catalogue of plants of the dendrological park “Sofiyivka”. Uman. (In Ukrainian)
Kosenko, I.S., & Mitin, V.V. (1995). The Uman State Dendrological Park “Sofiyivka” of the NAS of Ukraine as a center of introduction work in the Right-Bank Forest-Steppe of Ukraine. Introduction and Acclimatization of Plants, 23, 3–5. (In Russian)
Kostruba, T.M. (2020, July 6–9). Plants as symbols of national holidays. In Proceedings of the 3rd International Conference “Ethnobotanical traditions in agronomy, pharmacy and garden design” (pp. 174–180). Uman. (In Ukrainian)
Kostruba, T.M. (2024a). Annotated synopsis of decorative small-annual plants of the flora of the Middle Dnieper region. Journal of Native and Alien Plant Studies, 20, 62–76. (In Ukrainian). https://doi.org/10.37555/2707-3114.20.2024.318656
Kostruba, T.M. (2024b). Ephemeroids in urban and park plantings of the Uman region (Cherkasy Oblast). Biology and Ecology, 10(1), 26–31. (In Ukrainian). https://doi.org/10.33989/2024.10.1.306005
Kostruba, T.M. (2024c, September 30 – October 4). Naturalization of herbaceous ergasiophytes in the flora of the Middle Dnieper region. In Proceedings of the 15th Congress of the Ukrainian Botanical Society (p. 148). Helvetyka Publishing House, Odesa. (In Ukrainian)
Kostruba, T.M., & Chorna, H.A. (2021a, July 5–7). Introduction of ornamental plants in the Tsarytsyn Garden of Uman at the turn of the 19th and 20th centuries. In Materials of the 4th International Scientific Conference “Ethnobotanical traditions in agronomy, pharmacy, and garden design” (pp. 17–24). Lira-K, Kyiv. (In Ukrainian)
Kostruba, T.M., & Chorna, H.A. (2021b, March 31). Potentially invasive decorative species of Asteraceae in Ukraine. In Proceedings of the International Scientific Conference “Conservation of plants in connection with climate changes and biological invasions” (pp. 205–208). Bilotserkivdruk, Bila Tserkva. (In Ukrainian)
Kostruba, T.M., Chorna, G.A., & Mamchur, T.V. (2021). Thladiantha dubia Bunge as an invasively dangerous species in Ukraine. Journal of Native and Alien Plant Studies, 17(1), 183–188. (In Ukrainian). https://doi.org/10.37555/2707-3114.1.2021.247574
Kotov, M. (1928). Adventitious vegetation in Ukraine. Bulletin of Natural Studies, 5–6, 267–274. (In Ukrainian)
Kozlovska, N.A., Pashkevych, H.O., & Khamayko, N.V. (2013). Paleobotanical materials from excavations at Spaska Street, 35, Kyiv. Archaeology and Ancient History of Ukraine, 11, 90–99. (In Ukrainian)
Krytska, T.V., Levchuk, L.V., Chaban, K.V., Petrun, N.V., & Holokoz, A.V. (2010, October 11–15). Rare ornamental herbaceous plants in the collection of the Botanical Garden of Odesa National University. In Proceedings of the International Scientific Conference “The plant world in the Red Book of Ukraine: implementation of the global strategy for plant conservation” (pp. 278–282). Naukova Dumka, Kyiv. (In Ukrainian)
Kushnir, O. (2005). Decorative planting and flower composition features in cemeteries decoration. Plant Introduction, 25, 71–76. (In Ukrainian). https://doi.org/10.5281/zenodo.2586172
Leshcheniuk, E.N. (2014, May 20–24). The collection of rare ornamental perennials of the Kryvyi Rih Botanical Garden of the NAS of Ukraine as the basis for creation of culturophytocenoses of various purposes under conditions of Kryvorizhzhia. In Proceedings of the International Scientific Conference “Introduction, conservation and monitoring of biotic diversity” (pp. 74–75). Kyiv. (In Ukrainian)
Lunina, N.M. (2001). Cultural flora of ornamental herbaceous plants in Belarus and the role of introduction in increasing biodiversity. Bulletin of Poltava State Agricultural Institute, 1, 38–40. (In Russian)
Lunina, N.M., Volodko, I.K., Haishun, V.V., Svitkovska, O.I., & Ryzhenkova, Y.I. (2010). Ornamental herbaceous plants of the cultural flora of Belarus. Belaruskaya Navuka, Minsk. (In Russian)
Mamchur, T.V., Chorna, H.A., Parubok, M.I., Svystun, O.V., & Mykhailova, N.V. (2023). Catalogue of plants of the botanical nursery of Uman National University of Horticulture. Uman National University of Horticulture, Uman. (In Ukrainian)
Marynych, O.M., Parkhomenko, G.O., Petrenko, O.M. & Shishchenko, P.G. (2003). The improved scheme of the physical-geographical regionalization of Ukraine. Ukrainian Geographical Journal, 2, 16–20. (In Ukrainian)
Maryushkina, V. (2002). Adventization of vegetation as the consequence of spontaneous and purposeful plant introduction. Plant Introduction, 13, 49–60. (In Ukrainian). https://doi.org/10.5281/zenodo.3333777
Mashkovska, S.P. (Ed.). (2015). Catalogue of ornamental herbaceous plants of botanical gardens and arboretums of Ukraine. Electronic edition. Medobory, Kyiv. (In Ukrainian)
Mashkovska, S.P., & Pereboichuk, O.P. (2019). The collection fund of flowering and ornamental plants of the family Lamiaceae Martinov of M.M. Gryshko National Botanical Garden of the National Academy of Sciences of Ukraine. Plant Varieties Studying and Protection, 15(3), 249–258. (In Ukrainian). https://doi.org/10.21498/2518-1017.15.3.2019.181082
Mashtaler, N., Chipilyak, T., Mazura, M., & Bereslavskaya, O. (2011, June 8–11). Floral-decorative plants in greenery planting of Kryvyi Rih central part. In Proceedings of the 3rd International Conference on Landscape Architecture in Botanical Gardens and Arboretums (pp. 220–225). Kyiv. (In Ukrainian)
Mazur, T.P. (2000). Pond in the garden (Nymphaea – types, varieties, hybrids, features of development and care for them). Home Garden Backyard, Kyiv. (In Ukrainian)
Mazura, M.Y., Leshchenyuk, O.M., & Teslenko, I.K. (2020, September 22–24). Analysis of the range of herbaceous flower and ornamental plants of the Park “Feofaniya”. In Proceedings of the International Scientific Conference “Fundamental and applied aspects of plant introduction in the context of global environmental change” (pp. 131–134). Lira-K, Kyiv. (In Ukrainian)
Melnyk, V.I., Nesin, Y.D., & Shynder, O.I. (2015). Primula vulgaris (Primulaceae) – new species for the flora of Kyiv Polissya. Ukrainian Botanical Journal, 72(3), 241–245. (In Ukrainian). https://doi.org/10.15407/ukrbotj72.03.241
Michailovich, N. (2011). Phytosozological analysis of the decorative fraction of the flora of the National Natural Park “Skolivski Beskydy”. Florology and Phytosozology, 2, 245–247. (In Ukrainian)
Montresor, V. (1881). A review of the most beautiful plants that make up the flora of the provinces of the Kyiv educational district: Kyiv, Podolian, Volhynian, Chernihiv, and Poltava. Kyiv Society of Horticulture, Kyiv. (In Russian)
Montresor, V. (1898). List of plants collected in the Kyiv Educational Province in the last 25-year period. Zapiski Kievskogo Obshchestva Estestvoispytateley, 15(2), 675–707. (In Russian)
Moroz, I.I. (1970). Flora of the Tovtry Ridge in Podillia and its use in the national economy and for introduction (PhD thesis. Central Republican Botanical Garden of the Academy of Sciences of the Ukrainian SSR). Kyiv. (In Russian)
Moroz, I.I. (1983). Carnations of the natural flora for ornamental gardening. Naukova Dumka, Kyiv. (In Russian)
Mosyakin, S.L. (1991). Preliminary list of recent additions to the alien flora of Ukraine. Ukrainian Botanical Journal, 48(4), 28–34.
Mosyakin, S.L. (1996). Territorial peculiarities of expansion of alien plants in urbanized territories, with special reference to Kyiv, Ukraine. Ukrainian Botanical Journal, 53(5), 536–545. (In Ukrainian)
Mosyakin, S.L., & Mosyakin, A.S. (2021). Lockdown botany 2020: some noteworthy records of alien plants in Kyiv City and Kyiv Region. Ukrainian Botanical Journal, 78(2), 96–111. https://doi.org/10.15407/ukrbotj78.02.096
Mosyakin, S.L., & Yavorska, O.G. (2002). The nonnative flora of the Kyiv (Kyiv) urban area, Ukraine: a checklist and brief analysis. Urban Habitats, 1(1), 45–65.
Moysiyenko, I. (1997). Flora of cemeteries of Kherson. In Proceedings of the 10th Congress of the Ukrainian Botanical Society (pp. 1–192). Kyiv. (In Ukrainian)
Moysiyenko, I., Shynder, O., Levon, A., Chorna, G., Volutsa, O., Lavrinenko, K., Kolomiychuk, V., Shol, G., Shevera, M., Borovyk, D., Vynokurov, D., Zviahintseva, K., Kalashnik, K., Kazarinova, H., Levchuk, L., Skobel, H., Tarabun, M., Gerasimchuk, G., Lyubinska, L., Bezsmertna, O., Bondarenko, H., Mamchur, T., & Pashkevych, N. (2023). Notes to vascular plants in Ukraine. I. Chornomorski Botanical Journal. 19(1), 76–93. https://doi.org/10.32999/ksu1990-553X/2023-19-1-3
Muzychuk, G., & Pereboichuk, O. (2009). Ornamental plants of the genus Anemone L. in the world cultivated flora and the perspectives of their introduction in Ukraine. Plant Introduction, 44, 29–41. (In Ukrainian). https://doi.org/10.5281/zenodo.2555371
Muzychuk, G., & Prokopchuk, V. (2005). World assortment of cultivars of flower ornamental plants of Scrophulariaceae Juss. family and perspectives for their introduction into Ukraine. Plant Introduction, 25, 46–52. (In Ukrainian). https://doi.org/10.5281/zenodo.2586157
Nāburga, I., & Evarts-Bunders, P. (2019). Status of some escaped ornamental perennials in the flora of Latvia. Botanica, 25(2), 131–144. https://doi.org/10.2478/botlit-2019-0015
Naegeli, O., & Thellung, A. (1905). Die Flora des Kantons Zurich. Vierteljahrsschrift der Naturforschenden Gesellschaft in Zürich, 50, 225–305.
Nechitaylo, V.A., Badanina, V.A., & Gritsenko, V.V. (2005). The cultural plants of Ukraine. Phytosociocentre, Kyiv. (In Ukrainian)
Oksiiuk, P. (1924). On the issue of the spread of adventitious plants in Ukraine. Scientific Memoirs: Transactions of the Chairs for Scientific Researches in Kyiv, 2, 121–129. (In Ukrainian)
Onuk, L., Petruk, Y., & Chubata, T. (2021). The plant catalog of the phytosozology department. Tvory, Vinnytsia. (In Ukrainian)
Orlov, O. (2019, May 22–25). Wild ornamental adventive herbaceous plants and their invasion potential in Zhytomyr Oblast. In Materials of the International Scientific-Practical Conference “Introduction and preservation of plant diversity in botanical gardens of Eastern Europe” (pp. 54–61). Talcom, Kyiv. (In Ukrainian)
Orlov, O., Shynder, O., Chorna, H., Volutsa, O., Celka, Z., & Shevera, M. (2024). Potentilla indica (Rosaceae) in the flora of Ukraine: history of naturalization and modern distribution. Journal of Native and Alien Plant Studies, 20, 151–175. (In Ukrainian). https://doi.org/10.37555/2707-3114.20.2024.318679
Osadchyi, V.I., Kosovets, O.O., & Babichenko, V.M. (Eds.). (2010). Climate of Kyiv City. Nika Center, Kyiv. (In Ukrainian)
Paczoski, I. (1887). Essay on the flora of the environs of Uman, Kiev province. Notes of the Kiev Society of Naturalists, 8(2), 371–437. (In Russian)
Partytskiy, O.M. (1884). The tale of Ihor`s campaign [Slovo o polku Ihorevim]. Publishing House of the T.G. Shevchenko Scientific Society, Lviv. (In Old Ukrainian & Russian)
Pashkevych, H.A. (1991). Paleoethnobotanical finds on the territory of Ukraine. Ancient Rus: catalogue. Preprint, Kyiv. (In Russian)
Pavlenko, L.L. (2016). Ornamental herbaceous lianas in the Forest-Steppe of Ukraine: ontogenesis, reproductive capacity, use (PhD thesis. M.M. Gryshko National Botanical Garden of National Academy of Sciences of Ukraine). Kyiv. (In Ukrainian)
Pavlova, M.A., Kudina, G.A., & Kachur, L.Y. (2011). Decorative cereals of Pennisetum Rich. genus under the conditions of the southeast of Ukraine. Problems of Ecology and Nature Protection of the Technogenic Region, 1(11), 99–105. (In Ukrainian)
Pereboichuk, O.P. (2023, October 12). Prospects for the conservation and use of rare plants of the natural flora of Ukraine in landscape design. In Proceedings of the All-Ukrainian Scientific-Practical Conference “Theoretical and applied aspects of the study, conservation, and enrichment of plant diversity in research institutions and educational establishments of Ukraine” (pp. 151–154). PNU Korolenko, Poltava. (In Ukrainian)
Pereboichuk, O.P., Shcherbakova, T.O., & Mashkovska, S.P. (2021, March 12–13). Introduced plant species of the natural flora of Ukraine for ornamental gardening. In Proceedings of the International Scientific and Practical Conference “Ideas and innovations in natural sciences” (pp. 18–24). Baltija Publishing, Lublin. (In Ukrainian)
Philip, C., & Lord, T. (Eds.). (2003). RHS plant finder: 2003–2004. Dorling Kindersley Book.
Polonska-Vasylenko, N. (1993). History of Ukraine. Volume 1: until the mid-18th century. 2nd ed. Lybid, Kyiv. (In Ukrainian)
Popovych, S.Y., Vlasenko, A.S., & Vakarenko, O.V. (2018). Synopsis of decorative phytoautochthons of Ukraine. Komprint, Kyiv. (In Ukrainian)
POWO. (2025). Plants of the world online. Board of Trustees of the Royal Botanic Gardens, Kew. https://powo.science.kew.org/
Prokudin, Y.N., Vovk, A.H., Petrova, O.A., Ermolenko, E.D., & Vernichenko, Y.V. (1977). Grasses of Ukraine. Kyiv, Naukova Dumka. (In Russian)
Protopopova, V.V. (1988). Naturalization of adventitious plants of Ukraine. Ukrainian Botanical Journal, 45(4), 10–15. (In Ukrainian)
Protopopova, V.V. (1991). Synanthropic flora of Ukraine and ways of its development. Naukova Dumka, Kyiv. (In Russian)
Protopopova, V.V., & Shevera, M.V. (2013, September 10–12). Ergasiophytes as potential reserves of Ukrainian alien fraction flora. In Proceedings of the 1st International Scientific Conference “Non-traditional, new and forgotten plant species: scientific and practical aspects of cultivation” (pp. 99–101). Knyhonosha, Kyiv. (In Russian)
Protopopova, V.V., & Shevera, M.V. (2014). Ergasiophytes of the Ukrainian flora. Biodiversity Research and Conservation, 35(1), 31–46. https://doi.org/10.2478/biorc-2014-0018
Protopopova, V.V., & Shevera, M.V. (2019). Invasive species in the flora of Ukraine. I. Group of highly active species. GEO & BIO, 17, 116–135. (In Ukrainian). https://doi.org/10.15407/gb.2019.17.116
Protopopova, V.V., Mosyakin, S.L., & Shevera, M.V. (2002). Phytoinvasions in Ukraine as a threat to biodiversity: current state and future tasks. Kyiv. (In Ukrainian)
Pyšek, P., Richardson, D.M., Rejmánek, M., Webster, G.L., Williamson, M., & Kirscher, J. (2004). Alien plants in checklists and floras: towards better communication between taxonomists and ecologists. Taxon, 53(1), 131–143. https://doi.org/10.2307/4135498
Rakhmetov, D.B. (Ed.). (2015). Catalogue of plants of the department of new crops. Phytosociocentre, Kyiv. (In Ukrainian)
Rakhmetov, D.B., & Zaimenko, N.V. (2022). Stability of introduced and rare plants under climatic changes in Ukraine. Lira-K, Kyiv. (In Ukrainian)
Raunkiaer, C. (1907). Livsformernes Statistik som Grunlag for biologisk Plantegeografi. Botanisk Tidsskrift, 29, 42–83.
Richardson, D.M., Pyšek P., Rejmánek, M., Barbour, M.G., Panetta, F.D., & West, C.J. (2000). Naturalization and invasion of alien plants: concepts and definitions. Diversity and Distributions, 6(2), 93–107. https://doi.org/10.1046/j.1472-4642.2000.00083.x
Rikli, M. (1903). Die Antropochoren und der Formenkreis des Nasturtium palustre DC. Berichte der Schweizerischen Botanischen Gesellschaft, 13, 71–82.
Rogovich, A. (1855). An overview of vascular and semi-vascular plants that comprise the flora of the Kiev, Chernigov, and Poltava Governorates. Kyiv University Publishing House, Kyiv. (In Russian)
Rothmaler, W. (2017). Rothmaler – Excursionflora von Deutschland: Gefäßpflanzen. Springer Spektrum.
Rusanova, A.V., & Bengus, Y.V. (2020). Horticulture and ornamental landscaping as influential factors in the escape of invasive plants into natural environments. In Monitoring and Conservation of Biodiversity in Ukraine. Series: Conservation Biology in Ukraine, 16(1), 174–175. (In Ukrainian)
Schroeder, F.G. (1969). Zur Klassifizierung der anthropochoren. Vegetatio, 16(5/6), 225–238. https://doi.org/10.1007/BF00257018
Shabarova, S.I., Dyadyusha, L.M., & Shvets, I. (2002). Species composition of the decorative flora of homesteads in Kyiv and its environs. Scientific Bulletin of the National Agrarian University, 53, 302–305. (In Ukrainian)
Shcherbacova, T.O. (2014, May 20–24). Development and conservation of ornamental grasses collection in M.M. Gryshko National Botanical Gardens of the NAS of Ukraine. In Proceedings of the International Scientific Conference “Introduction, conservation and monitoring of biotic diversity” (p. 118). Kyiv. (In Ukrainian)
Shevera, M.V. (2017). Reynoutria × bohemica (Polygonaceae): a potentially invasive species in the flora of Ukraine. Ukrainian Botanical Journal, 74(6), 548–555. (In Ukrainian). https://doi.org/10.15407/ukrbotj74.06.548
Shiyan, N.M. (2011). Genus Paeonia L. (Paeoniaceae) in the Ukrainian flora. Ukrainian Botanical Journal, 68(1), 35–44. (In Ukrainian)
Shvets, T. (2006, September 25–28). Flowering and fruiting of species of the genus Iris L. as an indicator of successful introduction under cultural conditions. In Proceedings of the International Scientific Conference “Old parks and botanical gardens – scientific centers for biodiversity conservation and protection of historical and cultural heritage” (pp. 301–305). Uman. (In Ukrainian)
Shynder, O.I. (2010, September 15–17). Collection of taxa of Jovibarba Opiz and Sempervivum L. (Crassulaceae) in the M.M. Gryshko National Botanical Garden of NAS of Ukraine. In Proceedings of the International Scientific Conference “Introduction of plants, conservation, and enrichment of biodiversity in botanical gardens and dendrological parks” (pp. 347–349). Phytosociocenter, Kyiv. (In Ukrainian)
Shynder, O.I. (2019). Spontaneous flora of M.M. Gryshko National Botanical Garden of the NAS of Ukraine (Kyiv). 2. Methodological problems and criteria for selection of escaped plants in botanical garden conditions. Plant Introduction, 2, 3–16. (In Ukrainian). https://doi.org/10.5281/zenodo.3240995
Shynder, O.I. (2022). Cultivated plants of the Rzhyshchiv City amalgamated territorial community. In Studies of “Hlyboki Вalyky”. Ecological Research Station. Issue 2 (pp. 47–115). Druk Art, Chernivtsi. (In Ukrainian)
Shynder, O.I., & Negrash, Y.M. (2020). Sedum pallidum (Crassulaceae) – alien species of the flora of the plain part of Ukraine. Plant Introduction, 85/86, 75–84. https://doi.org/10.46341/PI2020009
Shynder, O.I., Bezsmertna, O.O., & Kucher, O.O. (2021). Flora of Rzhyshchiv City amalgamated territorial community: structure, regional features, synanthropic and rare species. In SA. Kuzemko, Y. Kutsokon, O. Vasilyuk (Eds.), Studies of “Hlyboki Вalyky” Ecological Research Station. Biodiversity of Rzhyshchiv City amalgamated territorial community. Issue 1 (pp. 15–100). Druk Art, Chernivtsi. (In Ukrainian)
Shynder, O.I., Doiko, N.M., Glukhova, S.A., Mykhajlyk, S.M., & Negrash, Y.M. (2022). New information about the flora of plant introduction institutions in Kyiv City and Bila Tserkva city (Kyiv Region). Chornomorski Botanical Journal, 18(1), 25–51. (In Ukrainian). https://doi.org/10.32999/ksu1990-553X/2022-18-1-2
Shynder, O.I., Nehrash, Y.M., Mamchur, T.V., & Kostruba, T.M. (2023a). Ornithogalum boucheanum (Asparagaceae) in Eastern Europe: native and synanthropic range, habitat conditions and state of population. Biosystems Diversity, 31(1): 59–70. https://doi.org/10.15421/012307
Shynder, O.I., Rak, O.O., Gritsenko, V.V., Nehrash, Y.M., Glukhova, S.A., Mykhailyk, S.M., & Kushnir, N.V. (2023b). Genus Corydalis s.l. (Papaveraceae) in the flora of Ukraine: wild and cultivated taxa and the success of naturalization of alien species. NaUKMA Research Papers. Biology and Ecology, 6, 48–59. (In Ukrainian). https://doi.org/10.18523/2617-4529.2023.6.48-59
Shynder, O.I., Davydov, D.A., Olshanskyi, I.G., Levon, A.F., & Nesyn, Y.D. (2024a). New floristic records in Kyiv City and its environs. Ukrainian Botanical Journal, 81(2), 100–144. (In Ukrainian). https://doi.org/10.15407/ukrbotj81.02.083
Shynder, O.I., Yatsentyuk, Y.V., Chorna, H.A., & Kostruba, T.M. (2024b). Flora of the “Synytsky Park” landscape gardening monument (Cherkasy Region). Chornomorski Botanical Journal, 20(4), 410–438. (In Ukrainian). https://doi.org/10.32999/ksu1990-553X/2024-20-4-4
Shynder, O.I., Kostruba, T.M., Chorna, H.A., & Mamchur, T.V. (2024c, September 11–12). Dynamics of the naturalization of herbaceous ergasiophytes in the botanical nursery of Uman National University of Horticulture (Cherkasy Region). In Proceedings of the 4th All-Ukrainian Scientific Conference “Synanthropization of the plant cover of Ukraine” (pp. 164–168). Bila Tserkva, Kyiv. (In Ukrainian)
Skybitska, M.I., & Mohyliak, M.H. (2003). Ornamental perennials in the medicinal plant collection of the Botanical Garden of Ivan Franko Lviv National University. Scientific Bulletin of Ukrainian National Forestry University, 13(5), 384–388. (In Ukrainian)
Slepchenko, L.O. (2001). Results of the introduction of ornamental plants in the “Askania-Nova” Dendrological Park. News of the Biosphere Reserve “Askania Nova”, 3, 38–41. (In Ukrainian)
Sobko, V.H., & Gaponenko, M.B. (1996). Introduction of rare and endangered plants of Ukraine’s flora. Naukova Dumka, Kyiv. (In Ukrainian)
Strgulc Krajšek, S., Bahčič, E., Čoko, U., & Dolenc Koce, J. (2020). Disposal methods for selected invasive plant species used as ornamental garden plants. Management of Biological Invasions, 11(2), 293–305. https://doi.org/10.3391/mbi.2020.11.2.08
Sydoruk, B.S. (1970). Ground cover and ornamental herbaceous plants of the natural flora of the “Sofiyivka” Dendrological Park. Introduction and Acclimatization of Plants in Ukraine, 1, 224–233. (In Ukrainian)
Sydoruk, T.N., & Sydoruk, B.S. (1992). Biology of some ground cover plants. Naukova Dumka, Kyiv. (In Russian)
Sytnik, K.M. (Ed.). (1984). Yarrows (Achillea). Naukova Dumka, Kyiv. (In Russian)
Tatarchuk, R., Kuznetsov, S., & Kazanskaya, N. (2012). Flowering-ornamental plants in mountain park and garden landscapes. Plant Introduction, 53, 114–119. (In Ukrainian). https://doi.org/10.5281/zenodo.2543918
Tzvelev, N.N. (Ed.). (1989–1994). Flora partis europaeae URSS. Vols. 7–8. Nauka, Leningrad. (In Russian)
Tzvelev, N.N. (Ed.). (1996–2004). Flora Europae Orientalis. Vols. 9–11. Mir i Semia, Petropoli. (In Russian)
Tzvelev, N.N., & Probatova, N.S. (2019). Grasses of Russia. KMK Scientific Press, Moscow. (In Russian)
UkrBin. (2025). Ukrainian Biodiversity Information Network. https://ukrbin.com
Vasylyeva, T.V., Nemertsalov, V.V., & Kovalenko, S.G. (2019a, September 25–27). Ornamental herbaceous plants of Odesa as an indicator of climate change. In Proceedings of the VII All-Ukrainian Congress of Ecologists with International Participation (Ecology–2019) (p. 139). Vinnytsia. (In Ukrainian).
Vasylyeva, T.V., Nemertsalov, V.V., & Kovalenko, S.G. (2019b). Synopsis of Odesa flora. Osvita Ukrainy, Odesa. (In Ukrainian)
von Raab-Straube, E., & Raus, T. (Eds.). (2024). Euro+Med-Checklist notulae, 17. Willdenowia, 54, 9–10. https://doi.org/10.3372/wi.54.54101
Vulf, E. V. (1987). Cultural flora of the globe: lists of taxa by floristic complexes. Nauka, Leningrad. (In Russian)
Vvedensky, A.I. (1935). Allium L. In V.L. Komarov (Ed.), Flora of the USSR. Vol. 4 (pp. 112–280). Academy of Sciences of USSR, Leningrad. (In Russian)
Vvedensky, A.I. (1971). Tulipa L. In A.I. Vvedenskij (Ed.), Determinant of the plants of Central Asia. Critical synopsis of flora. Vol. 2 (pp. 94–109). Fan, Tashkent. (In Russian)
Williamson, M. (1993). Invaders, weeds, and the risk from genetically modified organisms. Experientia, 49, 219–224. https://doi.org/10.1007/BF01923529
Williamson, M., & Fitter, A. (1996). The varying success of invaders. Ecology, 77(6), 1661–1666. https://doi.org/10.2307/2265769
Yanchuk, O., Smolinska, M., & Korolyuk, V. (2000). Ornamental herbaceous plants in green-belt settings of cities in Bucovyna. Plant Introduction, 5, 104–105. (In Ukrainian). https://doi.org/10.5281/zenodo.3338092
Yena, A.V. (2020). Lectures on ornamental plant growing: nomenclature of cultivated plants. Simferopol. (In Russian)
Zavialova, L.V. (2008). Aizopsis aizoon (L.) Grulich (Crassulaceae) – a new ergasiophyte in the flora of Ukraine. Ukrainian Botanical Journal, 65(6), 876–881. (In Ukrainian)
Zavialova, L.V. (2017). The most harmful invasive plant species for native phytodiversity of protected areas of Ukraine. Biological Systems, 9(1), 87–107. (In Ukrainian)
Zelinka, S.V., & Shymanska, V.O. (1976). Some ornamental plants of the Tovtry Ridge flora in Podillia and their protection. In Achievements of Botanical Science in Ukraine 1970–1973 (pp. 111–112). Naukova Dumka, Kyiv. (In Ukrainian)
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